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Adaptive evolution
He found that with enlarged clutches, more fledged and that their overall reproductive value had increased. However, even though the chicks in the larger clutches had increased fitness, their parents had actually suffered a decrease in fitness as they ended up having more responsibilities and less time to ensure their own survival.
This enlarged clutch size in the end decreased the residual reproductive value of the kestrels, therefore decreasing clutch size. So, the size at which the clutches were originally before Daan had tampered with them had evolved to be that size by means of natural selection as a compromise between fitness and fecundity
